In the distant future, Simon, a shy boy, and Kamina, a man who dreams of another life up on the surface live a quiet and restless life deep underground in Giha Village. One day, their destinies are forever changed when a gigantic “Gunmen” along with a beautiful girl named Yoko come falling through their village ceiling! Kamina, Simon, and Yoko break through to the surface riding the mysterious “Lagann” but the surface is nothing like Kamina imagined. Now, Kamina and Simon along with their comrades must challenge the evil Spiral King with the Gurren Lagann to bring hope to this desperate world!